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<!DOCTYPE article SYSTEM "http://www.clim-past.net/inc/cp/copernicus.dtd">
<article language="en">
	<journal>
		<journal_title>Climate of the Past</journal_title>
		<journal_url>www.clim-past.net</journal_url>
		<issn>1814-9324</issn>
		<eissn>1814-9332</eissn>
		<volume_number>5</volume_number>
		<issue_number>2</issue_number>
		<publication_year>2009</publication_year>
	</journal>
	<doi>10.5194/cp-5-203-2009</doi>
	<article_url>http://www.clim-past.net/5/203/2009/</article_url>
	<abstract_html>http://www.clim-past.net/5/203/2009/cp-5-203-2009.html</abstract_html>
	<fulltext_pdf>http://www.clim-past.net/5/203/2009/cp-5-203-2009.pdf</fulltext_pdf>
	<start_page>203</start_page>
	<end_page>216</end_page>
	<publication_date>2009-06-03</publication_date>
	<article_title content_type="html">Late Quaternary vegetation-climate feedbacks</article_title>
	<authors>
		<author numeration="1" affiliations="1,2,3">
			<name>M. Claussen*</name>
		</author>
	</authors>
	<affiliations>
		<affiliation numeration="1" content_type="html">Max Planck Institute for Meteorology, Bundesstr. 53, 20146 Hamburg, Germany</affiliation>
		<affiliation numeration="2" content_type="html">Meteorological Institute, University Hamburg, Bundesstr. 55, 20146 Hamburg, Germany</affiliation>
		<affiliation numeration="3" content_type="html">Invited contribution by M. Claussen, recipient of the EGU Milutin Milankovic Medal 2005.</affiliation>
	</affiliations>
	<abstract content_type="html">Feedbacks between vegetation and other components of the climate system are
discussed with respect to their influence on climate dynamics during the
late Quaternary, i.e., the last glacial-interglacial cycles.
When weighting current understanding based on interpretation of palaeobotanic and
palaeoclimatic evidence by numerical climate system models, a number of
arguments speak in favour of vegetation dynamics being an amplifier of
orbital forcing. (a) The vegetation-snow albedo feedback in synergy with
the sea-ice albedo feedback tends to amplify Northern Hemisphere and global
mean temperature changes. (b) Variations in the extent of the largest desert
on Earth, the Sahara, appear to be amplified by biogeophysical feedback. (c)
Biogeochemical feedbacks in the climate system in relation to vegetation
migration are supposed to be negative on time scales of glacial cycles.
However, with respect to changes in global mean temperature, they are
presumably weaker than the positive biogeophysical feedbacks.</abstract>
	<references>
		<reference numeration="1" content_type="text"> % vor jede Referenz Anhuf, D., Frankenberg, P., and Lauer, W.: Die postglaziale Warmphase vor 8000 Jahren, Geol. Rundsch. 51, 454â€“461, 1999. </reference>
		<reference numeration="2" content_type="text"> % vor jede Referenz Archer, D., Winguth, A., Lea, D., and Mahowald, N.: What caused the glacial/interglacial atmospheric $p$CO&lt;sub&gt;2&lt;/sub&gt; cycles? Rev. Geophys., 38, 159â€“189, 2000. </reference>
		<reference numeration="3" content_type="text"> % vor jede Referenz Bala, G., Caldeira, K., Wickett, M., Phillips, T. J., Lobell, D. B., Delire, C., and Mirin, A.: Combined climate and carbon-cycle effects of large-scale deforestation, 17 April, PNAS, 104, 6550â€“6555, 2007. </reference>
		<reference numeration="4" content_type="text"> % vor jede Referenz Berger, A.: Long-term variations of daily insolation and quaternary climatic change, Journal of Atmospheric and Oceanic Science, 35, 2362â€“2367, 1978. </reference>
		<reference numeration="5" content_type="text"> % vor jede Referenz Berger, A.: The role of CO&lt;sub&gt;2&lt;/sub&gt;, sea-level and vegetation during the Milankovitch-forced glacial-interglacial cycles, in: Geosphere-Biosphere Interactions and Climate, edited by: Bengtsson, L. O. and Hammer, C. U., Vatican City, Cambridge University Press, New York, 119â€“146, 2001. </reference>
		<reference numeration="6" content_type="text"> % vor jede Referenz Betts, R. A.: Offset of the potential carbon sink from boreal forestation by decreases in surface albedo, Nature, 408, 187â€“190, 2000. </reference>
		<reference numeration="7" content_type="text"> % vor jede Referenz Bonan, G. B.: Forests and Climate Change: Forcings, Feedbacks, and the Climate Benefits of Forests, Science, 320, 1444â€“1449, 2008. </reference>
		<reference numeration="8" content_type="text"> % vor jede Referenz Braconnot, P., Joussaume, S., Marti, O., and deNoblet-Ducoudre, N.: Synergistic feedbacks from ocean and vegetation on the African monsoon response to mid-Holocene insolation, Geophys. Res. Lett., 26(16), 2481â€“2484, 1999. </reference>
		<reference numeration="9" content_type="text"> % vor jede Referenz Braconnot, P., Otto-Bliesner, B., Harrison, S., Joussaume, S., Peterchmitt, J.-Y., Abe-Ouchi, A., Crucifix, M., Driesschaert, E., Fichefet, Th., Hewitt, C. D., Kageyama, M., Kitoh, A., Laîné, A., Loutre, M.-F., Marti, O., Merkel, U., Ramstein, G., Valdes, P., Weber, S. L., Yu, Y., and Zhao, Y.: Results of PMIP2 coupled simulations of the Mid-Holocene and Last Glacial Maximum â€“ Part 1: experiments and large-scale features, Clim. Past, 3, 261â€“277, 2007. </reference>
		<reference numeration="10" content_type="text"> % vor jede Referenz Brovkin, V., Claussen, M., Petoukhov, V., and Ganopolski, A.: On the stability of the atmosphere-vegetation system in the Sahara/Sahel region, J. Geophys. Res., 103(D24), 31613â€“31624, 1998. </reference>
		<reference numeration="11" content_type="text"> % vor jede Referenz Brovkin, V., Bendtsen, J., Claussen, M., Ganopolski, A., Kubatzki, C., Petoukhov, V., and Andreev, A.: Carbon cycle, vegetation and climate dynamics in the Holocene: Experiments with the CLIMBER-2 model, Global Biogeochem. Cy., 16(4), 1139, doi:10.1029/2001GB001662, 2002. </reference>
		<reference numeration="12" content_type="text"> % vor jede Referenz Brovkin, V., Levis, S., Loutre, M. F., Crucifix, M., Claussen, M., Ganopolski, A., Kubatzki, C., and Petoukhov, V.: Stability analysis of the climate-vegetation system in the northern high latitudes, Climatic Change, 57(1), 119â€“138, 2003. </reference>
		<reference numeration="13" content_type="text"> % vor jede Referenz Brovkin, V., Ganopolski, A., Archer, D., and Rahmstorf, S.: Lowering of glacial $p$CO&lt;sub&gt;2&lt;/sub&gt; in response to changes in oceanic circulation and marine biogeochemistry, Paleoceanography, 22, PA4202, doi:10.1029/2006PA001380, 2007. </reference>
		<reference numeration="14" content_type="text"> % vor jede Referenz BrostrÃ¶m, A., Coe, M., Harrison, S. P., Gallimore, R., Kutzbach, J. E., Foley, J. A., Prentice, I. C., and Behling, P.: Land surface feedbacks and palaeomonsoons in northern Africa, Geophys. Res. Lett., 25(19), 3615â€“3618, 1998. </reference>
		<reference numeration="15" content_type="text"> % vor jede Referenz Chang, P., Zhang, R., Hazeleger, W., Wen, C., Wan, X., Ji, L., Haarsma, R. J., Breugem, W.-P., and Seidel, H.: Oceanic link between abrupt changes in the North Atlantic Ocean and the African monsoon, Nature Geoscience, 1(7), 444â€“448, 2008. </reference>
		<reference numeration="16" content_type="text"> % vor jede Referenz Charney, J. G.: Dynamics of deserts and droughts in the Sahel, Q. J. Roy. Meteorol. Soc., 101, 193â€“202, 1975. </reference>
		<reference numeration="17" content_type="text"> % vor jede Referenz Cheddadi, R., Yu, G., Guiot, J., Harrison, S. P., and Prentice, I. C.: The climate of Europe 6000 years ago, Clim. Dyn., 13, 1â€“9, 1997. </reference>
		<reference numeration="18" content_type="text"> % vor jede Referenz Claussen, M.: On coupling global biome models with climate models, Climate Res., 4, 203â€“221, 1994. </reference>
		<reference numeration="19" content_type="text"> % vor jede Referenz Claussen, M.: Modelling biogeophysical feedback in the African and Indian Monsoon region, Clim. Dynam., 13, 247â€“257, 1997. </reference>
		<reference numeration="20" content_type="text"> % vor jede Referenz Claussen, M. and Gayler, V.: The greening of Sahara during the mid-Holocene: Results of an interactive atmosphere-biome model, Global Ecol. Biogeogr., 6, 369â€“377, 1997. </reference>
		<reference numeration="21" content_type="text"> % vor jede Referenz Claussen, M., Kubatzki, C., Brovkin, V., Ganopolski, A., Hoelzmann, P., and Pachur, H. J.: Simulation of an abrupt change in Saharan vegetation at the end of the mid-Holocene, Geophys. Res. Lett., 24, 2037â€“2040, 1999. </reference>
		<reference numeration="22" content_type="text"> % vor jede Referenz Claussen, M., Brovkin, V., Petoukhov, V., and Ganopolski, A.: Biogeophysical versus biogeochemical feedbacks of large-scale land-cover change, Geophys. Res. Letters, 26(6), 1011â€“1014, 2001. </reference>
		<reference numeration="23" content_type="text"> % vor jede Referenz Claussen, M., Ganopolski, A., Brovkin, V., Gerstengarbe, F.-W., and Werner, P.: Simulated Global-Scale Response of the Climate System to Dansgaard-Oeschger and Heinrich events, Clim. Dynam., 21, 361â€“370, 2003a. </reference>
		<reference numeration="24" content_type="text"> % vor jede Referenz Claussen, M., Brovkin, V., Ganopolski, A., Kubatzki, C., and Petoukhov, V.: Climate Change in Northern Africa: The past is not the future, Climatic Change, 57(1), 99â€“118, 2003b. </reference>
		<reference numeration="25" content_type="text"> % vor jede Referenz Claussen, M. (Ed.): Does land surface matter in climate and weather?, in: Vegetation, Water, Humans and the Climate, Part A., edited by: Kabat, P., Claussen, M., Dirmeyer, P., Gash, J., de Guenni, L. B., Meybeck, M., Pielke Sr., R. A., VÃ¶rÃ¶smarty, C., Hutjes, R., and LÃ¼tkemeier, S., Springer-Verlag, Heidelberg, 5â€“153, 2004. </reference>
		<reference numeration="26" content_type="text"> % vor jede Referenz Claussen, M., Cox, P. M., Zeng, X., Viterbo, P., Beljaars, A. C. M., Betts, R., Bolle, H.-J., Chase, T., and Koster, R.: The Global Climate â€“ Chapter A.4., in: Vegetation, Water, humans and the climate: A new perspective on an interactive system, edited by: Kabat, P., Claussen, M., Dirmeyer, P. A., Gash, J. H. C., de Guenni, L. B., Meybeck, M. Pielke Sr., R. A., VÃ¶rÃ¶smarty, C. J., and LÃ¼tkemeier, S., Springer-Verlag, Heidelberg, 33â€“57, 2004. </reference>
		<reference numeration="27" content_type="text"> % vor jede Referenz Claussen, M., Fohlmeister, J., Ganopolski, A., and Brovkin, V.: Vegetation dynamics amplifies precessional forcing, Geophys. Res. Lett., 33, L09709, doi:10.1029/2006GL026111, 2006. </reference>
		<reference numeration="28" content_type="text"> % vor jede Referenz Coe, M. T. and Harrison, S. P.: The water balance of northern Africa during the mid-Holocene: An evaluation of the 6 ka BP PMIP simulations, Clim. Dynam., 19, 155â€“166, 2002. </reference>
		<reference numeration="29" content_type="text"> % vor jede Referenz Cox, P. M., Betts, R. A., Jones, C. D., Spall, S. A., and Totterdell, I. J.: Acceleration of global warming due to carbon-cycle feedbacks in a coupled climate model, Nature, 404, 184â€“187, 2000. </reference>
		<reference numeration="30" content_type="text"> % vor jede Referenz Cremaschi, M., Pelfini, M., and Santilli, M.: Cupressus dupreziana: a dendroclimatic record for the middle-late Holocene in the central Sahara, The Holocene, 16(2), 293â€“303, 2006. </reference>
		<reference numeration="31" content_type="text"> % vor jede Referenz Crowley, T. J.: Ice age terrestrial carbon changes revisited, Global Biogeochem. Cy., 9(3), 377â€“389, 1995. </reference>
		<reference numeration="32" content_type="text"> % vor jede Referenz Crucifix, M. and Loutre, M. F.: Transient simulations over the last interglacial period (126â€“115 kyr BP): feedback and forcing analysis, Clim. Dynam, 19(5â€“6), 417â€“433, 2002. </reference>
		<reference numeration="33" content_type="text"> % vor jede Referenz Crucifix, M., Loutre, M. F., Tulkens, P., Fichefet, T., and Berger, A.: Climate evolution during the Holocene: a study with an Earth system model of intermediate complexity, Clim. Dynam, 19(1), 43â€“60, 2002. </reference>
		<reference numeration="34" content_type="text"> % vor jede Referenz deMenocal, P. B., Ortiz, J., Guilderson, T., Adkins, J., Sarnthein, M., Baker, L., and Yarusinski, M.: Abrupt onset and termination of the African Humid Period: Rapid climate response to gradual insolation forcing, Quarternary Sci. Rev., 19, 347â€“361, 2000. </reference>
		<reference numeration="35" content_type="text"> % vor jede Referenz Denman, K. L., Brasseur, G., Chidthaisong, A., Ciais, P., Cox, P. M., Dickinson, R. E., Hauglustaine, D., Heinze, C., Holland, E., Jacob, D., Lohmann, U., Ramachandran, S., da Silva Dias, P. L., Wofsy, S. C. and Zang, X.: Couplings between changes in the climate system and Biogeochemistry, in: Climate Change 2007: The Physical Science Basis, Contribution of Working Group I to the Fourth Assessment Report of the Intergovernmental Panel on Climate Change, edited by: Solomon, S., Qin, D., Manning, M., Chen, Z., Marquis, M., Averyt, K. B., Tignor, M., and Miller, H. L., Cambridge University Press, Cambridge, UK and New York, NY, USA, 499â€“587, 2007. </reference>
		<reference numeration="36" content_type="text"> % vor jede Referenz de Noblet, N., Prentice, I. C., Joussaume, S., Texier, D., Botta, A., and Haxeltine, A.: Possible role of atmosphere-biosphere interactions in triggering the last glaciation, Geophys. Res. Lett., 23(22), 3191â€“3194, 1996. </reference>
		<reference numeration="37" content_type="text"> % vor jede Referenz Doherty, R., Kutzbach, J., Foley, J., and Pollard, D.: Fully coupled climate/dynamical vegetation model simulations over Northern Africa during the mid-Holocene, Clim. Dynam., 16, 561â€“573, 2000. </reference>
		<reference numeration="38" content_type="text"> % vor jede Referenz Elenga, H., Peyron, O., Bonnefille, R., Jolly, D., Cheddadi, R., Guiot, J., Andrieu, V., Bottema, S., Buchet, G., De Beaulieu, J.-L., Hamilton, A. C., Maley, J., Marchant, R., Perez-Obiol, R., Reille, M., Riollet, G., Scott, L., Straka, H., Taylor, D., Van Campo, E., Vincens, A., Laarif, F., and Jonson, H.: Pollen-based biome reconstruction for southern Europe and Africa 18 000 yr bp, J. Biogeogr., 27, 621â€“634, 2000. </reference>
		<reference numeration="39" content_type="text"> % vor jede Referenz Eltahir, E. A. B. and Gong, C.: Dynamics of wet and dry years in West Africa, J. Climate, 9(5), 1030â€“1042, 1996. </reference>
		<reference numeration="40" content_type="text"> % vor jede Referenz Foley, J. A., Kutzbach, J. E., Coe, M. T., and Levis, S.: Feedbacks between climate and boreal forests during the Holocene epoch, Nature, 371, 52â€“54, 1994. </reference>
		<reference numeration="41" content_type="text"> % vor jede Referenz Foley, J. A.: Tipping points in the Tundra, Science, 310, 627â€“628, 2005. </reference>
		<reference numeration="42" content_type="text"> % vor jede Referenz Francois, L. M., Godderis, Y., Warnant, P., Ramstein, G., de Noblet, N., and Lorenz, S.: Carbon stocks and isotopic budgets of the terrestrial biosphere at mid-Holocene and last glacial maximum times, Chem. Geol., 159, 163â€“189, 1999. </reference>
		<reference numeration="43" content_type="text"> % vor jede Referenz Friedlingstein, P., Bopp, L., Ciais, P., Dufresne, J.-L., Fairhead, L., LeTreut, H., Monfray, P., and Orr, J.: Positive feedback between future climate change and the carbon cycle, Geophys. Res. Lett., 28(8), 1543â€“1546, 2001. </reference>
		<reference numeration="44" content_type="text"> % vor jede Referenz Friedlingstein, P., Cox, P., Betts, R., Bopp, L., von Bloh, W., Brovkin, V., Cadule, P., Doney, S., Eby, M., Fung, I., Bala, G., John, J., Jones, C., Joos, F., Kato, T., Kawamiya, M., Knorr, W., Lindsay, K., Matthews, H. D., Raddatz, T., Rayner, P., Reick, C., Roeckner, E., Schnitzler, K.-G., Schnur, R., Strassmann, K., Weaver, A. J., Yoshikawa, C., and Zeng, N.: Climate-carbon cycle feedback analysis, results from the C4MIP model intercomparison, J. Climate, 19, 3337â€“3353, 2006. </reference>
		<reference numeration="45" content_type="text"> % vor jede Referenz Frenzel, B., Pesci, M., and Velichko, A. A. (Eds.): Atlas of paleoclimates and paleoenvironments of the Northern Hemisphere: Late Pleistocene â€“ Holocene, Budapest: Geographical Research Institute, 153 pp., 1992. </reference>
		<reference numeration="46" content_type="text"> % vor jede Referenz Gallimore, R. G., Jacob J., and Kutzbach, J. E.: Coupled atmosphere-ocean-vegetation simulations for modern and mid-Holocene climates: role of extratropical vegetation cover feedbacks, Clim. Dynam., 25, 755â€“776, 2006. </reference>
		<reference numeration="47" content_type="text"> % vor jede Referenz Ganopolski, A., Kubatzki, C., Claussen, M., Brovkin, V., and Petoukhov, V.: The influence of vegetation-atmosphere-ocean interaction on climate during the mid-Holocene, Science, 280, 1916â€“1919, 1998. </reference>
		<reference numeration="48" content_type="text"> % vor jede Referenz Ganopolski, A.: Glacial Integrative modelling, Philos. T. R. Soc. A, London, 361, 1871â€“1884, 2003. </reference>
		<reference numeration="49" content_type="text"> % vor jede Referenz Grassl, H.: Status and improvements of coupled general circulation models, Science, 288, 1991â€“1997, 2000. </reference>
		<reference numeration="50" content_type="text"> % vor jede Referenz Hewitt, C. D. and Mitchell, J. F. B.: A fully coupled GCM simulation of the climate of the mid-Holocene, Geophys. Res. Lett., 25(3), 361â€“364, 1998. </reference>
		<reference numeration="51" content_type="text"> % vor jede Referenz Hoelzmann, P., Jolly, D., Harrison, S. P., Laarif, F., Bonnefille, R., and Pachur, H.-J.: Mid-Holocene land-surface conditions in northern Africa and the Arabian Peninsula: A data set for the analysis of biogeophysical feedbacks in the climate system, Global Biogeochem. Cy., 12, 35â€“51, 1998. </reference>
		<reference numeration="52" content_type="text"> % vor jede Referenz House, J. I., Brovkin, V., Betts, R., Constanza, R., Silva Dias, M. A., Holland, E., le Quere, C., Kim Phat, N., Riebesell, U., and Scholes, M.: Climate and air quality, in Ecosystems and Human Well-Being: Current State and Trends, Findings of the Condition and Trends Working Group of the Millennium Ecosystem Assessment, Island Press, Washington, Covelo, London, 1, 357â€“390, 2006. </reference>
		<reference numeration="53" content_type="text"> % vor jede Referenz Irizarry-Ortiz, M., Wang, G., and Elthair, E. A. B.: Role of the biosphere in the mid-Holocene climate of West Africa, J. Geophys. Res., 108(D2), 4042, doi:10.1029/2001JD000989, 2003. </reference>
		<reference numeration="54" content_type="text"> % vor jede Referenz Jahn, A.: Atmosphere-vegetation feedback in the climate system of the last glacial, Diploma thesis, Fachbereich Geowissenschaften, Freie UniversitÃ¤t Berlin, Germany, 2004. </reference>
		<reference numeration="55" content_type="text"> % vor jede Referenz Jahn, A., Claussen, M., Ganopolski, A., and Brovkin, V.: Quantifying the effect of vegetation dynamics on the climate of the Last Glacial Maximum, Clim. Past, 1, 1â€“7, 2005. </reference>
		<reference numeration="56" content_type="text"> % vor jede Referenz Jolly, D., Prentice, I. C., Bonnefille, R., et al.: Biome reconstruction from pollen and plant macrofossil data for Africa and Arabian Peninsula at 0 and 6 k, J. Biogeogr., 25, 1007â€“1027, 1998. </reference>
		<reference numeration="57" content_type="text"> % vor jede Referenz Joos, F., Gerber, S., Prentice, I. C., Otto-Bliesner, B. L., and Valdes, P. J.: Transient simulations of Holocene atmospheric carbon dioxide and terrestrial carbon since the Last Glacial Maximum, Global Biogeochem. Cy., 18, GB2002, doi:10.1029/2003GB002156, 2004. </reference>
		<reference numeration="58" content_type="text"> % vor jede Referenz Joussaume, S., Taylor, K. E., Braconnot, P., Mitchell, J. F. B., Kutzbach, J. E., Harrison, S. P., Prentice, I. C., Broccoli, A. J., Abe-Ouchi, A., Bartlein, P. J., Bonfiels, C., Dong., B., Guiot, J., Herterich, K., Hewit, C. D., Jolly, D., Kim, J. W., Kislov, A., Kitoh, A., Loutre, M. F., Masson, V., McAvaney, B., McFarlane, N., deNoblet, N., Peltier, W. R., Peterschmitt, J. Y., Pollard, D., Rind, D., Royer, J. F., Schlesinger, M. E., Syktus, J., Thompson, S., Valdes, P., Vettoretti, G., Webb, R. S., and Wyputta, U.: Monsoon changes for 6000 years ago: Results of 18 simulations from the Paleoclimate Modeling Intercomparison Project (PMIP), Geophys. Res. Lett., 26(7), 859â€“862, 1999. </reference>
		<reference numeration="59" content_type="text"> % vor jede Referenz Kageyama, M., Charbit, S., Ritz, C., Khodri, M., and Ramstein, G.: Mechanisms leading to the last glacial inception over North America.: Results from the CLIMBER-GREMLINS atmosphere-ocean-vegetation northern hemisphere ice-sheet, model, in: The climate of past interglacials, edited by: Sirocko, F., Claussen, M., Sanchez Goni, M. F., and Litt, T., Elsevier, Amsterdam, 573â€“582, 2007. </reference>
		<reference numeration="60" content_type="text"> % vor jede Referenz Kirchner, J. W.: The GAIA Hypothesis: can it be tested? Rev. Geophys., 27(2), 223â€“235, 1989. </reference>
		<reference numeration="61" content_type="text"> % vor jede Referenz Knorr, W. and Schnitzler, K.-G.: Enhanced albedo feedback in North Africa from possible combined vegetation and soil formation processes, Clim. Dynam., 26, 55â€“63, 2006. </reference>
		<reference numeration="62" content_type="text"> % vor jede Referenz KÃ¶ppen, W. and Wegener, A.: Die Klimate der geologischen Vorzeit, Borntraeger, Berlin, 255 pp., 1924. </reference>
		<reference numeration="63" content_type="text"> % vor jede Referenz KrÃ¶pelin, S., Verschuren, D., LÃ©zine, A.-M., Eggermont, H., Cocquyt, C., Francus, P., Cazet, J.-P., Fagot, M., Russell, J. M., Darius, F., Conley, D. J., Schuster, M., von Suchodoletz, H., and Engstrom, D. R.: Climate-driven ecosystem succession in the Sahara: The past 6000 years, Science, 30, 765â€“768, 2008. </reference>
		<reference numeration="64" content_type="text"> % vor jede Referenz Kubatzki, C. and Claussen, M.: Simulation of the global biogeophysical interactions during the last glacial maximum, Clim. Dynam., 14, 461â€“471, 1998. </reference>
		<reference numeration="65" content_type="text"> % vor jede Referenz Kubatzki, C., Montoya, M., Rahmstorf, S., Ganopolski, A., and Claussen, M.: Comparison of a coupled global model of intermediate complexity and an AOGCM for the last interglacial, Clim. Dynam., 16, 799â€“814, 2000. </reference>
		<reference numeration="66" content_type="text"> % vor jede Referenz Kubatzki, C., Claussen, M., Calov, R., and Ganopolski, A.: Sensitivity of the last glacial inception to initial and surface conditions, Clim. Dynam., 27, 333â€“344, 2006. </reference>
		<reference numeration="67" content_type="text"> % vor jede Referenz Kubatzki, C., Claussen, M., Calov, R., and Ganopolski, A.: Modelling the end of an interglacial (MIS 1,5,7,9,11), in: The climate of past interglacials, edited by: Sirocko, F., Claussen, M., Litt, T., and Sanchez-Goni, M., Elsevier, Amsterdam, 583â€“593, 2007. </reference>
		<reference numeration="68" content_type="text"> % vor jede Referenz Kutzbach, J. E. and Guetter, P. J.: The influence of changing orbital parameters and surface boundary conditions on climate simulations for the past 18 000 years, J. Atmos. Sci., 43, 1726â€“1759, 1986. </reference>
		<reference numeration="69" content_type="text"> % vor jede Referenz Kutzbach, J. E., Gallimore, R., Harrison, S., Behling, P., Selin, R., and Laarif, F.: Climate and biome simulations for the past 21 000 years, Quarternary Sci. Rev., Elsevier Science Ltd., 17, 473â€“506, 1998. </reference>
		<reference numeration="70" content_type="text"> % vor jede Referenz Kutzbach, J. E. and Liu, Z.: Oceanic feedback on the western African monsoon at 6000 BP, Science, 278, 440â€“443, 1997. </reference>
		<reference numeration="71" content_type="text"> % vor jede Referenz Levis, S., Foley, J. A., Brovkin, V., and Pollard, D.: On the stability of the high-latitude climate-vegetation system in a coupled atmosphere-biosphere model, Global Ecol. Biogeogr., 8, 489â€“500, 1999. </reference>
		<reference numeration="72" content_type="text"> % vor jede Referenz Lovelock, J. E. and Margulis, L.: Atmospheric hoeostasis by and for the biosphere: the gaia hypothesis, Tellus, 26, 1â€“9, 1974. </reference>
		<reference numeration="73" content_type="text"> % vor jede Referenz Liu, Z., Harrison, S. P., Kutzbach, J. E., and Otto-Bliesner, B.: Global monsoons in the mid-Holocene and oceanic feedback, Clim. Dynam., 22, 157â€“182, 2004. </reference>
		<reference numeration="74" content_type="text"> % vor jede Referenz Liu, Z., Notaro, M., Kutzbach J., and Liu, N.: Assessing global vegetation-climate feedbacks from the observation, J. Climate, 19(5), 787â€“814, 2006a. </reference>
		<reference numeration="75" content_type="text"> % vor jede Referenz Liu, Z., Wang, Y., Gallimore, R., Notaro, M., and Prentice, I. C.: On the cause of abrupt vegetation collapse in North Africa during the Holocene: Climate variability vs. vegetation feedback, Geophys. Res. Lett., 33, L22709, doi:10.1029/2006GL028062, 2006b. </reference>
		<reference numeration="76" content_type="text"> % vor jede Referenz Liu, Z., Wang, Y., Gallimore, R., Gasse, F., Johnson, T., deMenocal, P., Adkins, J., Notaro, M., Prentice, I. C., Kutzbach, J. E., Jacob, R., Behling, P., Wang, L., and Ong, E.: Simulating the transient evolution and abrupt change of Northern Africa Atmosphere-Ocean-Terrestrial Ecosystem in the Holocene, Quarternary Sci. Rev., 26, 1818â€“1837, 2007. </reference>
		<reference numeration="77" content_type="text"> % vor jede Referenz MacDonald, G. M., Velichko, A. A., Kremenetski, C. V., Borisova, O. K., Goleva, A. A., Andreev, A. A., Cwynar, L. C., Riding, R. T., Forman, S. L., Edwards, T. W. D., Aravena, R., Hammarlund, D., Szeicz, J. M., and Gattaulin, V. N.: Holocene treeline history and climate change across Northern Eurasia, Quaternary Res., 53, 302â€“311, 2000. </reference>
		<reference numeration="78" content_type="text"> % vor jede Referenz Meehl, G. A., Stocker, T. F., Collins, W. D., Friedlingstein, P., Gaye, A. T., Gregory, J. M., Kitoh, A., Knutti, R., Murphy, J. M., Noda, A., Raper, S. C. B., Watterson, I. G., Weaver, A. J., and Zhao, Z.-C.: Global Climate Projections, in: Climate Change 2007: The Physical Science Basis Contribution of Working Group I to the Fourth Assessment Report of the Intergovernmental Panel on Climate Change, edited by: Solomon, S., Qin, D., Manning, M., Chen, Z., Marquis, M., Averyt, K. B., Tignor, M., and Miller H. L., Cambridge University Press, Cambridge, UK and New York, NY, USA, 747â€“845, 2007. </reference>
		<reference numeration="79" content_type="text"> % vor jede Referenz Meissner, K. J., Weaver, A. J., Matthews, H. D., and Cox, P. J.: The role of land-surface dynamics in glacial inception: a study with the UVic Earth System Model, Clim. Dynam., 21, 515â€“537, 2003. </reference>
		<reference numeration="80" content_type="text"> % vor jede Referenz Notaro, M., Wang, Y., Liu, L., Gallimore, R., and Levis, S.: Combined statistical and dynamical assessment of the simulated negative vegetation feedback on North African annual precipitation during the mid-Holocene, Global Change Biol., 14, 347â€“368, 2008. </reference>
		<reference numeration="81" content_type="text"> % vor jede Referenz Osborne, A. H., Vance, D., Rohling, E. J., Barton, N., Rogerson, M., and Fello, N.: A humid corridor across the Sahara for the migration of early modern humans out of Africa 120 000 years ago, PNAS, 105, 16444â€“16447, 2008. </reference>
		<reference numeration="82" content_type="text"> % vor jede Referenz Otterman, J.: Baring high-albedo soils by overgrazing: A hypothesized desertification mechanism, Science, 186, 531â€“533, 1974. </reference>
		<reference numeration="83" content_type="text"> % vor jede Referenz Otterman, J., Chou, M.-D., and Arking, A.: Effects of nontropical forest cover on climate, J. Clim. Appl. Meteorol., 23, 762â€“767, 1984. </reference>
		<reference numeration="84" content_type="text"> % vor jede Referenz Otto, J., Raddatz, T., Claussen, M., Brovkin, V., and Gayler, V.: Separation of atmosphere-ocean-vegetation feedbacks and synergies for mid-Holocene climate, Geophys. Res. Lett., 36, L09701, doi:10.1029/209GL037482, 2009. </reference>
		<reference numeration="85" content_type="text"> % vor jede Referenz Petit-Maire, N. and Guo, Z.: Mise en Ã©vidence de variations climatiques holocÃ¨nes rapides, en phase dans les dÃ©serts actuels de Chine et du Nord de l&apos;Afrique, Sciences de la Terre et des PlanÃ¨tes, 322, 847â€“851, 1996. </reference>
		<reference numeration="86" content_type="text"> % vor jede Referenz Pachur, H.-J.: PalÃ¤o-Environment und Drainagesysteme der Ostsahara im SpÃ¤tpleistozÃ¤n und HolozÃ¤n, in: Nordost-Afrika: Strukturen und Ressourcen; Ergebnisse aus dem Sonderforschungsbereich &quot;Geowissenschaftliche Probleme in Ariden und Semiariden Gebieten&quot;, edited by: Klitzsch, E. and Thorweihe, U. , Wiley â€“ VCH, Weinheim, 366â€“455, 1999. </reference>
		<reference numeration="87" content_type="text"> % vor jede Referenz Prentice, I. C., Sykes, M. T., Lautenschlager, M., Harrison, S. P., Denissenko, O., and Bartlein, P.: Modelling global vegetation patterns and terrestrial carbon storage at the last glacial maximum, Global Ecol. Biogeogr., 3, 67â€“76, 1993. </reference>
		<reference numeration="88" content_type="text"> % vor jede Referenz Prentice, I. C. and Webb III, T.: BIOME 6000: reconstructing global mid-Holocene vegetation patterns from palaeoecological records, J. Biogeogr., 25, 997â€“1005, 1998. </reference>
		<reference numeration="89" content_type="text"> % vor jede Referenz Prentice, I. C., Jolly, D., and BIOME 6000 members: Mid-Holocene and glacial-maximum vegetation geography of the northern continents and Africa, J. Biogeogr., 27, 507â€“519, 2000. </reference>
		<reference numeration="90" content_type="text"> % vor jede Referenz Raddatz, T. J., Reick, C. H., Knorr, W., Kattge, J., Roeckner, E., Schnur, R., Schnitzler, K.-G., Wetzel, P., and Jungclaus, J.: Will the tropical land biosphere dominate the climate-carbon cycle feedback during the twenty-first century? Clim. Dynam., 29, 565â€“574, 2007. </reference>
		<reference numeration="91" content_type="text"> % vor jede Referenz Renssen, H., Brovkin, V., Fichefet, T., and Goosse, H.: Holocene climate instability during the termination of the African Humid Period, Geophys. Res. Lett., 30(4), 1184, doi:10.1029/2002/GL016636, 2003. </reference>
		<reference numeration="92" content_type="text"> % vor jede Referenz Scheffer, M., Carpenter, S., Foley, J. A., Folke, C., and Walker, B.: Catastrophic shifts in ecosystems, Nature, 413, 591â€“596, 2001. </reference>
		<reference numeration="93" content_type="text"> % vor jede Referenz Scheffer, M., Brovkin, V., and Cox, P. M.: Positive feedback between global warming and atmospheric CO&lt;sub&gt;2&lt;/sub&gt; concentration inferred from past climate change, Geophys. Res. Lett., 33, L10702, doi:10.1029/2005GL025044, 2006. </reference>
		<reference numeration="94" content_type="text"> % vor jede Referenz Schurgers, G., Mikolajewicz, U., GrÃ¶ger, M., Maier-Reimer, E., VizcaÃ­no, M., and Winguth, A.: Dynamics of the terrestrial biosphere, climate and atmospheric CO&lt;sub&gt;2&lt;/sub&gt; concentration during interglacials: a comparison between Eemian and Holocene, Clim. Past, 2, 205â€“220, 2006. </reference>
		<reference numeration="95" content_type="text"> % vor jede Referenz Spitaler, R.: Das Klima des Eiszeitalters, Selbstverlag, Prag, 138 pp., 1921. </reference>
		<reference numeration="96" content_type="text"> % vor jede Referenz Stein, U. and Alpert, P.: Factor separation in numerical simulations, J. Atmos. Sci., 50, 2107â€“2115, 1993. </reference>
		<reference numeration="97" content_type="text"> % vor jede Referenz Tarasov, P. E., Webb III, T., Andreev, A. A., Afanaseva, N. B., Berezina, N. A., Bezusko, L. G., Blyakharchuk, T. A., Bolikhovskaya, N. S., Cheddadi, R., Chernavskaya, M. M., Chernova, G. M., Dorofeyuk, N. I., Dirksen, V. G., Elina, G. A., Filimonova, L. V., Glebov, F. Z., Guiot, J., Gunova, V. S., Harrison, S. P., Jolly, D., Khomutova, V. I., Kvavadze, E. V., Osipova, I. M., Panova, N. K., Prentice, I. C., Saarse, L., Sevastyanov, D. V., Volkova, V. S., and Zernitskaya, V. P.: Present-day and mid-Holocene biomes reconstructed from pollen and plant macrofossil data from the former Soviet Union and Mongolia, J. Biogeogr., 25, 1029â€“1053, 1998. </reference>
		<reference numeration="98" content_type="text"> % vor jede Referenz TEMPO: Potential role of vegetation feedback in the climate sensitivity of high-latitude regions: A case study at 6000 years B.P., Global Biogeochem. Cy., 10(4), 727â€“736, 1996. </reference>
		<reference numeration="99" content_type="text"> % vor jede Referenz Texier, D., de Noblet, N., Harrison, S. P., Haxeltine, A., Jolly, D., Joussaume, S., Laarif, F., Prentice, I. C., and Tarasov, P.: Quantifying the role of biosphere-atmosphere feed-backs in climate change: coupled model simulations for 6000 years BP and comparison with palaeodata for northern Eurasia and northern Africa, Clim. Dynam., 13, 865â€“882, 1997. </reference>
		<reference numeration="100" content_type="text"> % vor jede Referenz Tjallingii, R., Claussen, M., Stuut, J.-B. W., Fohlmeister, J., Jahn, A., Bickert, T., Lamy, F., and RÃ¶hl, U.: Coherent high- and low latitude control of the Northwest African hydrological balance and continental vegetation cover, Nature Geoscience, 1, 670â€“675, 2008. </reference>
		<reference numeration="101" content_type="text"> % vor jede Referenz Wang, Y., Notaro, M., Liu, Z., Gallimore, R., Levis, S., and Kutzbach, J. E.: Detecting vegetation-precipitation feedbacks in mid-Holocene North Africa from two climate models, Clim. Past, 4, 59â€“67, 2008. </reference>
		<reference numeration="102" content_type="text"> % vor jede Referenz Wang, Y., Mysak, L .A., Wang, Z., and Brovkin, V.: The greening of the McGill Paleoclimate Model. Part I: Improved land surface scheme with vegetation dynamics, Clim. Dynam., 24, 469â€“480, 2005a. </reference>
		<reference numeration="103" content_type="text"> % vor jede Referenz Wang, Y., Mysak, L. A., Wang, Z., and Brovkin, V.: The greening of the McGill Paleoclimate Model. Part II: Simulation of Holocene millennial-scale natural climate changes, Clim. Dynam., 24, 481â€“496, 2005b. </reference>
		<reference numeration="104" content_type="text"> % vor jede Referenz Wang, Z. M., Cochelin, A. S. B., Mysak, L. A., and Wang, Y.: Simulation of the last glacial inception with the green McGill Paleoclimate Model, Geophys. Res. Lett., 32(12), L12705, doi:10.1029/2005GL023047, 2005. </reference>
		<reference numeration="105" content_type="text"> % vor jede Referenz Watson, A. J. and Lovelock, J. E.: Biological Homeostasis of the Global Environment: The Parable of Daisyworld, Tellus, 35B, 284â€“289, 1983. </reference>
		<reference numeration="106" content_type="text"> % vor jede Referenz Weber, S. L.: On homeostasis in Daisyworld, Climatic Change, 48, 465â€“485, 2001. </reference>
		<reference numeration="107" content_type="text"> % vor jede Referenz Wohlfahrt, J., Harrison, S. P., and Braconnot, P.: Synergetic feedbacks between ocean and vegetation on mid- and high-latitude climates during the mid-Holocene, Clim Dynam., 22, 223â€“238, 2004. </reference>
		<reference numeration="108" content_type="text"> % vor jede Referenz Xue, Y. and Shukla, J.: The influence of land surface properties on Sahel climate. Part I: Desertification, J. Climate, 6, 2232â€“2245, 1993. </reference>
		<reference numeration="109" content_type="text"> % vor jede Referenz Yu, G. and Harrison, S. P.: An evaluation of the simulated water balance of Eurasia and northern Africa at 6000 y BP using lake status data, Clim. Dynam., 12, 723â€“735, 1996. </reference>
		<reference numeration="110" content_type="text"> % vor jede Referenz Zeng, N.: Glacial-interglacial atmospheric CO&lt;sub&gt;2&lt;/sub&gt; change â€“ The glacial burial hypothesis, Adv. Atmos. Sci., 20(5), 677â€“693, 2003. </reference>
	</references>
</article>

